Filoha Hamadryas Project Publications

Amann, A., Pines, M., & Swedell, L. (2017) Contexts and consequences of takeovers in hamadryas baboons: female parity, reproductive state, and observational evidence of pregnancy loss. American Journal of Primatology 79:e22649 (DOI: 10.1002/ajp.22649).

Städele, V., Pines, M., Swedell, L., & Vigilant, L. (2016) The ties that bind: maternal kin bias in a multilevel primate society despite natal dispersal by both sexes. American Journal of Primatology (DOI:10.1002/ajp.22537).

Swedell, L. & Chowdhury, S. (2016) Hamadryas Baboon: Papio hamadryas. Invited contribution to All the World’s Primates, edited by N. Rowe & M. Myers. Charlestown, RI: Pogonias Press, pp. 436-437.

Chowdhury, S., Pines, M., Saunders, J., & Swedell, L. (2015) The adaptive value of secondary males in the polygynous multi-level society of hamadryas baboons. Americal Journal of Physical Anthropology 158: 501-513 (DOI: 10.1002/ajpa.22804).

Pines, M., Chowdhury, S., Saunders, J., & Swedell, L. (2015) The rise and fall of leader males in a multi-level society: takeovers and tenures of male hamadryas baboons. American Journal of Primatology 77: 44-55 (DOI: 10.1002/ajp.22309).

Städele, V., Van Doren, V., Pines, M., Swedell, L., & Vigilant, L. (2015) Fine-scale genetic assessment of sex-specific dispersal patterns in a multilevel primate society. Journal of Human Evolution 78: 103-113 (

Swedell, L., Leedom, L., Saunders, J., & Pines, M. (2014) Sexual Conflict in a Polygynous Primate: Costs and Benefits of a Male-Imposed Mating System. Behavioral Ecology and Sociobiology 68: 263-273 (

Schreier, A. & Grove, M. (2014) Recurrent Patterning in the Daily Foraging Routes of Hamadryas Baboons (Papio hamadryas): Spatial Memory in Large‐Scale Versus Small‐Scale Space. American Journal of Primatology 76: 421–435 (

Swedell, L. & Plummer, T. (2012) A Papionin Multilevel Society as a Model for Hominin Social Evolution. International Journal of Primatology 33: 1165-1193 (

Schreier, A. & Swedell, L. (2012) The Socioecology of Network Scaling Ratios in the Multilevel Society of Hamadryas Baboons. International Journal of Primatology 33: 1069-1080 (DOI: 10.1007/s10764-011-9572-1).

Matsuda, I., Zhang, P., Swedell, L., Mori, U., Tuuga, A., Bernard, H., & Sueur, C. (2012) Comparisons of Inter-Individual Relationships among Nonhuman Primates Living in Multilevel Societies. International Journal of Primatology 33: 1038-1053 (DOI: 10.1007/s10764-012-9616-1).

Schreier, A., & Swedell, L. (2012) Ecology and Sociality in a Multilevel Society: Ecological Determinants of Social Cohesion in Hamadryas Baboons. American Journal of Physical Anthropology 148: 580-588 (DOI: 10.1002/ajpa.22076).

Pines, M., Saunders, J., & Swedell, L. (2011) Alternative Routes to the Leader Male Role: Follower versus Solitary Male Strategies and Outcomes in Hamadryas Baboons.  American Journal of Primatology 73: 679-691 (DOI: 10.1002/ajp.20951).

Pines, M. & Swedell, L.  (2011) Not without a Fair Fight: Failed Abductions of Females in Wild Hamadryas Baboons.  Primates 52: 249-252 (DOI: 10.1007/s10329-011-0242-x).

Swedell, L., Saunders, J., Schreier, A., Davis, B., Tesfaye, T., & Pines, M. (2011) Female “Dispersal” in Hamadryas Baboons: Transfer among Social Units in a Multilevel Society. American Journal of Physical Anthropology 145: 360-370 (DOI: 10.1002/ajpa.21504).

Schreier, A. & Grove, M. (2010) Ranging Patterns of Hamadryas Baboons: Random Walk Analyses. Animal Behaviour 80: 75-87.

Schreier, A. (2010) Feeding Ecology, Food Availability and Ranging Patterns of Wild Hamadryas Baboons at Filoha. Folia Primatologica 81: 129-145.

Schreier, A. & Swedell, L. (2009) The Fourth Level of Social Structure in a Multi-Level Society: Ecological and Social Functions of Clans in Hamadryas Baboons. American Journal of Primatology 71: 948-955.

Swedell, L. & Schreier, A. (2009) Male Aggression Towards Females in Hamadryas Baboons: Conditioning, Coercion, and Control. IN Sexual Coercion in Primates: An Evolutionary Perspective on Male Aggression Against Females, edited by M. Muller and R. Wrangham.  Cambridge: Harvard University Press, pp. 244-268.

Swedell, L., Hailemeskel, G., & Schreier, A. (2008) Composition and Seasonality of Diet in Wild Hamadryas Baboons: Preliminary Findings from Filoha. Folia Primatologica 79: 476-490.

Schreier, A. & Swedell, L. (2008) Use of Palm Trees as a Sleeping Site by Hamadryas Baboons in Ethiopia.  American Journal of Primatology 70: 107-113.

Swedell, L. & Saunders, J. (2006) Infant Mortality, Paternity Certainty, and Female Reproductive Strategies in Hamadryas Baboons. IN Reproduction and Fitness in Baboons: Behavioral, Ecological, and Life History Perspectives, edited by L. Swedell & S. Leigh.  New York: Springer, pp. 19-51.

Swedell, L. (2006) Strategies of Sex and Survival in Hamadryas Baboons: Through a Female Lens, Pearson Prentice Hall, Upper Saddle River, NJ.

Swedell, L. & Tesfaye, T. (2003) Infant Mortality After Takeovers in Wild Ethiopian Hamadryas Baboons. American Journal of Primatology 60: 113-118.

Swedell, L. (2002a) Affiliation Among Females in Wild Hamadryas Baboons (Papio hamadryas hamadryas). International Journal of Primatology 23: 1205-1226 (

Swedell, L. (2002b) Ranging Behavior, Group Size, and Behavioral Flexibility in Ethiopian Hamadryas Baboons (Papio hamadryas hamadryas).  Folia Primatologica 73: 95-103.

Swedell, L. (2000) Two Takeovers in Wild Hamadryas Baboons. Folia Primatologica 71: 169-172.